玉米育種特點及選育
一、玉米育種的特點
yumizuizhuyaodetezhengshitianranyihuachuanfen,tianranshoufenquntidetianjianzuchengchuyugaodudeyizhizhuangtai,getidejiyinxingchuyugaodudezahezhuangtai,zhejuedinglezaiyumitianranshoufendequntizhong,zhujianbiaoxianxingbijiaoyiyibuda,bixutongguoyidingdejiyinxingxuanzeguochengcainengzhengquedijuedingqushe;同tong時shi,由you於yu個ge體ti基ji因yin型xing高gao度du雜za合he,造zao成cheng表biao型xing選xuan擇ze不bu可ke靠kao,必bi須xu對dui大da量liang個ge體ti做zuo測ce交jiao或huo後hou代dai鑒jian定ding,才cai能neng確que認ren表biao型xing是shi否fou真zhen實shi遺yi傳chuan。由you於yu這zhe些xie原yuan因yin,在zai玉yu米mi育yu種zhong過guo程cheng中zhong,一yi般ban都dou要yao經jing過guo多duo代dai的de選xuan擇ze比bi較jiao才cai能neng育yu成cheng新xin的de自zi交jiao係xi或huo品pin種zhong。
現xian代dai玉yu米mi育yu種zhong的de主zhu流liu是shi雜za種zhong優you勢shi育yu種zhong,基ji本ben途tu徑jing是shi先xian選xuan育yu純chun合he的de親qin本ben自zi交jiao係xi,再zai將jiang親qin本ben自zi交jiao係xi雜za交jiao,選xuan育yu出chu雜za種zhong優you勢shi強qiang的de雜za交jiao種zhong。生sheng產chan上shang利li用yong的de是shiF1代dai的de雜za種zhong優you勢shi。自zi交jiao係xi的de選xuan育yu不bu但dan要yao求qiu本ben身shen性xing狀zhuang優you良liang,還hai要yao求qiu配pei合he力li高gao。對dui自zi交jiao係xi農nong藝yi性xing狀zhuang和he配pei合he力li的de選xuan擇ze具ju有you同tong等deng重zhong要yao的de意yi義yi,不bu可ke偏pian廢fei。這zhe就jiu大da大da提ti高gao了le育yu種zhong的de難nan度du,延yan長chang了le育yu種zhong的de周zhou期qi。不bu僅jin如ru此ci,自zi交jiao係xi的de性xing狀zhuang同tong雜za交jiao種zhong的de性xing狀zhuang雖sui然ran有you關guan係xi,但dan仍reng然ran有you距ju離li,在zai雜za交jiao種zhong水shui平ping上shang還hai要yao對dui農nong藝yi性xing狀zhuang進jin行xing選xuan擇ze鑒jian定ding,同tong時shi對dui主zhu要yao目mu標biao性xing狀zhuang的de雜za種zhong優you勢shi水shui平ping進jin行xing比bi較jiao,最zui終zhong育yu成cheng優you良liang雜za交jiao品pin種zhong。
二、玉米自交係的選育
1.農藝性狀好。植株性狀:主要包括株型和抗倒性。植株性狀一般根據穗上部葉片伸展的姿態分為緊湊型、半緊湊型、和平展型;根據株高分為高稈、中稈、矮稈、半矮稈等。
穗部性狀:穗部性狀一般由穗型、粒型、穗行數等構成。穗型的劃分有很多種,最明顯的是長穗型、粗穗型、筒型、錐型、紡錘型等;粒型分為馬齒型和硬粒型及其各種過渡類型。自交係選育中最好兼顧長穗型和粗穗型的選擇。長穗型的行粒數較多,但粒行數較少;粗穗型的粒行數較多,但行粒數不可能很多。自交係的粒行數一般10~20行。12~14xingdebijiaoshizhong。lixingdexuanzeyaogenjuyuzhongmubiaoqueding,yibanpianyinglixingdezijiaoxizupeidezajiaozhongshangpinxinghao,pianmachirongyizupeichudezajiaozhongchanlianghedianfenhanliangbijiaogao。ciwai,zilidedaxiao、粒重和粒色的一致性也需要考慮。
抗逆性:對主要的玉米病害和自然災害性條件要有一定的抗性或耐受性,以確保種子生產的穩定性和雜種優勢的穩定發揮。整齊一致性:要求農藝性狀在外觀上要整齊一致,體現在基因型上要基本純合。
2.配pei合he力li高gao。配pei合he力li是shi指zhi雜za交jiao親qin本ben在zai其qi雜za種zhong後hou代dai的de雜za種zhong優you勢shi中zhong發fa揮hui作zuo用yong的de潛qian在zai能neng力li。自zi交jiao係xi配pei合he力li的de高gao低di決jue定ding著zhe未wei來lai育yu成cheng雜za交jiao種zhong的de增zeng產chan能neng力li和he利li用yong價jia值zhi,因yin而er是shi自zi交jiao係xi最zui重zhong要yao的de性xing狀zhuang。配pei合he力li分fen為wei一yi般ban配pei合he力li和he特te殊shu配pei合he力li,前qian者zhe的de遺yi傳chuan基ji礎chu是shi來lai源yuan於yu親qin本ben的de基ji因yin加jia性xing效xiao應ying,後hou者zhe決jue定ding於yu親qin本ben基ji因yin的de非fei加jia性xing效xiao應ying,二er者zhe在zai遺yi傳chuan上shang具ju有you相xiang對dui的de獨du立li性xing。所suo以yi,在zai自zi交jiao係xi的de選xuan育yu上shang二er者zhe不bu可ke偏pian廢fei。
3.種子生產性能強。高產、優質、kangxinghaodezajiaozhongnengfoutuiguanghuozhechengweizhuzaipinzhong,henzhongyaodetiaojianshiyaojubeiyouliangdezhongzishengchanxingneng。baokuoqinbenzhongzifayalihefayashiqiang,youmiaochangshihao,yiyubaomiao,cixiongxietiao;父本散粉好,花粉量大,花期較長;母本雄穗柄長,抽雄容易,吐絲快而整齊,結實性好,籽粒均勻,產量高;父母本光溫反應小,花期協調容易,製種易於成功。
4.選xuan育yu自zi交jiao係xi的de基ji礎chu材cai料liao。從cong根gen本ben上shang說shuo,玉yu米mi雜za交jiao種zhong強qiang大da的de雜za種zhong優you勢shi來lai源yuan於yu親qin本ben間jian充chong分fen大da的de遺yi傳chuan差cha異yi。因yin此ci,選xuan育yu自zi交jiao係xi的de基ji礎chu材cai料liao必bi須xu來lai源yuan多duo樣yang,遺yi傳chuan基ji礎chu廣guang泛fan。選xuan係xi材cai料liao的de來lai源yuan大da體ti有you4個方麵:第一種是地方種質,包括老的地方品種群體,或者來源於地方品種的自交係等。在一些特殊的生態條件下可能存在優良抗源、或(huo)優(you)良(liang)品(pin)質(zhi)材(cai)料(liao),應(ying)該(gai)特(te)別(bie)注(zhu)意(yi)收(shou)集(ji)。第(di)二(er)種(zhong)是(shi)雜(za)交(jiao)種(zhong),主(zhu)要(yao)指(zhi)生(sheng)產(chan)上(shang)應(ying)用(yong)的(de)或(huo)者(zhe)引(yin)進(jin)的(de)雜(za)交(jiao)種(zhong),也(ye)包(bao)括(kuo)用(yong)自(zi)交(jiao)係(xi)有(you)目(mu)的(de)組(zu)配(pei)的(de)用(yong)於(yu)選(xuan)係(xi)的(de)雜(za)交(jiao)種(zhong),其(qi)中(zhong)應(ying)用(yong)最(zui)大(da)的(de)是(shi)單(dan)交(jiao)種(zhong)。一(yi)般(ban)認(ren)為(wei),采(cai)用(yong)當(dang)地(di)或(huo)異(yi)地(di)生(sheng)產(chan)上(shang)廣(guang)為(wei)利(li)用(yong)的(de)雜(za)交(jiao)種(zhong)選(xuan)係(xi)是(shi)育(yu)成(cheng)優(you)良(liang)自(zi)交(jiao)係(xi)的(de)捷(jie)徑(jing),這(zhe)些(xie)選(xuan)係(xi)一(yi)般(ban)農(nong)藝(yi)性(xing)狀(zhuang)好(hao),配(pei)合(he)力(li)比(bi)較(jiao)高(gao)。第(di)三(san)種(zhong)是(shi)綜(zong)合(he)種(zhong)或(huo)輪(lun)回(hui)改(gai)良(liang)的(de)群(qun)體(ti),這(zhe)些(xie)群(qun)體(ti)一(yi)般(ban)優(you)良(liang)基(ji)因(yin)的(de)頻(pin)率(lv)和(he)優(you)良(liang)基(ji)因(yin)型(xing)的(de)重(zhong)組(zu)率(lv)都(dou)比(bi)較(jiao)高(gao),但(dan)有(you)利(li)基(ji)因(yin)的(de)聚(ju)合(he)程(cheng)度(du)可(ke)能(neng)不(bu)如(ru)單(dan)交(jiao)種(zhong),在(zai)選(xuan)係(xi)過(guo)程(cheng)中(zhong)通(tong)常(chang)要(yao)加(jia)大(da)選(xuan)擇(ze)量(liang)。第(di)四(si)種(zhong)是(shi)遠(yuan)緣(yuan)雜(za)種(zhong)後(hou)代(dai)。玉(yu)米(mi)的(de)野(ye)生(sheng)近(jin)緣(yuan)種(zhong)很(hen)多(duo),其(qi)中(zhong)含(han)有(you)豐(feng)富(fu)的(de)優(you)良(liang)性(xing)狀(zhuang),但(dan)由(you)於(yu)種(zhong)間(jian)雜(za)交(jiao)困(kun)難(nan),後(hou)代(dai)不(bu)育(yu)或(huo)分(fen)離(li)嚴(yan)重(zhong),穩(wen)定(ding)慢(man),直(zhi)接(jie)用(yong)於(yu)選(xuan)育(yu)自(zi)交(jiao)係(xi)難(nan)度(du)大(da),現(xian)在(zai)已(yi)經(jing)很(hen)少(shao)使(shi)用(yong)。
5.選育自交係的方法。
(1)常chang規gui選xuan育yu法fa。這zhe種zhong選xuan育yu方fang法fa就jiu是shi在zai一yi個ge分fen離li的de原yuan始shi群qun體ti中zhong選xuan擇ze個ge體ti自zi交jiao係xi,經jing若ruo幹gan世shi代dai按an目mu測ce自zi交jiao選xuan擇ze之zhi後hou,進jin行xing配pei合he力li的de測ce定ding,最zui終zhong選xuan出chu優you良liang的de自zi交jiao係xi。常chang規gui選xuan係xi法fa的de問wen題ti是shi早zao期qi入ru選xuan的de規gui模mo很hen難nan界jie定ding。目mu前qian,有you的de育yu種zhong者zhe在zai早zao期qi世shi代dai不bu分fen穗sui行xing種zhong植zhi,而er是shi將jiang入ru選xuan的de果guo穗sui的de等deng量liang混hun合he種zhong植zhi,到dao晚wan代dai才cai嚴yan格ge選xuan擇ze單dan株zhu種zhong成cheng穗sui行xing,進jin行xing配pei合he力li測ce定ding。這zhe種zhong方fang法fa有you效xiao地di增zeng加jia了le入ru選xuan基ji因yin型xing的de數shu目mu,同tong時shi增zeng加jia了le對dui單dan株zhu的de選xuan擇ze壓ya力li,可ke能neng對dui提ti高gao抗kang性xing和he獲huo得de目mu標biao基ji因yin型xing有you幫bang助zhu。在zai育yu種zhong規gui模mo不bu大da的de情qing況kuang下xia,常chang規gui選xuan育yu法fa,更geng多duo地di依yi靠kao育yu種zhong者zhe的de經jing驗yan。
(2)單倍體選係法。其基本原理是利用自然或人工的誘發、培pei育yu的de單dan倍bei體ti植zhi株zhu經jing過guo人ren工gong的de染ran色se體ti加jia倍bei或huo自zi然ran加jia倍bei獲huo得de純chun合he的de二er倍bei體ti,然ran後hou再zai從cong中zhong選xuan育yu優you良liang的de單dan株zhu,成cheng為wei自zi交jiao係xi。單dan倍bei體ti育yu種zhong一yi般ban隻zhi需xu兩liang年nian就jiu能neng獲huo得de純chun合he的de自zi交jiao係xi,比bi常chang規gui方fang法fa縮suo短duan了le育yu種zhong的de周zhou期qi,對dui育yu種zhong家jia有you很hen大da的de吸xi引yin力li。不bu過guo,單dan倍bei體ti選xuan育yu省sheng略lve了le大da田tian選xuan擇ze的de步bu驟zhou,對dui諸zhu如ru抗kang性xing等deng性xing狀zhuang通tong常chang還hai要yao進jin行xing加jia倍bei的de選xuan擇ze。
(3)輻(fu)射(she)與(yu)化(hua)學(xue)誘(you)變(bian)選(xuan)係(xi)法(fa)。輻(fu)射(she)誘(you)變(bian)選(xuan)係(xi)法(fa)是(shi)利(li)用(yong)放(fang)射(she)性(xing)同(tong)位(wei)素(su)產(chan)生(sheng)的(de)輻(fu)射(she)能(neng)量(liang)甚(shen)至(zhi)宇(yu)宙(zhou)射(she)線(xian)誘(you)發(fa)染(ran)色(se)體(ti)產(chan)生(sheng)結(jie)構(gou)變(bian)異(yi)或(huo)等(deng)位(wei)基(ji)因(yin)突(tu)變(bian),根(gen)據(ju)表(biao)型(xing)變(bian)異(yi)選(xuan)擇(ze)有(you)用(yong)的(de)個(ge)體(ti)育(yu)成(cheng)自(zi)交(jiao)係(xi)。此(ci)種(zhong)方(fang)法(fa)的(de)誘(you)變(bian)機(ji)理(li)主(zhu)要(yao)以(yi)損(sun)傷(shang)恢(hui)複(fu),在(zai)恢(hui)複(fu)過(guo)程(cheng)中(zhong)發(fa)現(xian)有(you)利(li)的(de)變(bian)異(yi),因(yin)而(er)在(zai)選(xuan)擇(ze)原(yuan)始(shi)材(cai)料(liao)上(shang)要(yao)兼(jian)顧(gu)遺(yi)傳(chuan)基(ji)礎(chu)豐(feng)富(fu)和(he)抗(kang)損(sun)傷(shang)能(neng)力(li)強(qiang),在(zai)選(xuan)係(xi)過(guo)程(cheng)中(zhong)要(yao)注(zhu)意(yi)淘(tao)汰(tai)各(ge)種(zhong)畸(ji)形(xing)株(zhu)和(he)不(bu)利(li)變(bian)異(yi),選(xuan)擇(ze)突(tu)變(bian)性(xing)狀(zhuang)明(ming)顯(xian)的(de)健(jian)康(kang)的(de)植(zhi)株(zhu)。這(zhe)種(zhong)方(fang)法(fa)近(jin)年(nian)來(lai)被(bei)有(you)些(xie)人(ren)移(yi)植(zhi)為(wei)航(hang)天(tian)育(yu)種(zhong)或(huo)太(tai)空(kong)育(yu)種(zhong)。
(4)分子選係法。所謂分子選係法是在DNA水shui平ping上shang開kai展zhan的de自zi交jiao係xi選xuan育yu,包bao括kuo轉zhuan基ji因yin選xuan係xi法fa和he分fen子zi標biao記ji輔fu助zhu選xuan係xi法fa。轉zhuan基ji因yin選xuan係xi就jiu是shi借jie助zhu遺yi傳chuan轉zhuan化hua技ji術shu,將jiang一yi些xie其qi他ta動dong植zhi物wu有you利li的de外wai源yuan基ji因yin導dao入ru自zi交jiao係xi或huo雜za交jiao種zhong內nei,育yu成cheng有you外wai源yuan基ji因yin控kong製zhi性xing狀zhuang的de優you良liang自zi交jiao係xi,這zhe種zhong方fang法fa對dui於yu改gai造zao玉yu米mi基ji因yin庫ku中zhong不bu具ju備bei的de某mou些xie目mu標biao性xing狀zhuang是shi致zhi關guan重zhong要yao的de。分fen子zi標biao記ji輔fu助zhu選xuan係xi是shi首shou先xian對dui某mou些xie優you良liang基ji因yin確que定ding的de緊jin密mi連lian鎖suo的de分fen子zi標biao記ji,然ran後hou以yi這zhe些xie分fen子zi標biao記ji為wei間jian接jie選xuan擇ze性xing狀zhuang對dui目mu標biao性xing狀zhuang進jin行xing選xuan擇ze,最zui後hou育yu成cheng優you良liang自zi交jiao係xi。
(5)zijiaoxidegailiang。mouxiezijiaoxizaiyuchenghoufaxianyouxiebixukefudequedian,youxiezijiaoxizaiyingyongyiduanshijianhoufaxianmouxiezhongyaodexingzhuangfashengtuihua,huofashenglexindebinghaishenglixiaozhongdeqinran,huozhebushiyingxindetuiguangquyudehuanjing,zheshijiejuewentidejiejingshiduiyuanyouzijiaoxidegebiexingzhuangjinxinggaizao,yuchengxindegailiangxi。
改良自交係最常用的方法是回交轉育法。回交轉育法就是以被改良的自交係為輪回親本,以目的基因供體親本為非輪回親本,經過5代以上的回交改良和1~2daidezijiaowendinghou,jiukeyiyuchengxindegailiangzijiaoxi。huijiaoxiaoguodehaohuaitongmubiaoxingzhuangdeyichuanfangshiyouguan,yibanhuijiaofaduijiandanxingzhuanghuodanjiyinkongzhidexingzhuangyijixibaozhiyichuanxingzhuangzuiyouxiao,duifuzaxingzhuangxiaoguocha。huijiaofazaixuanzegongtiqinbenshiyaozhuyizhuyaodemubiaoxingzhuangbuyaoyouliansuoleizhui,haiyaozhuyixuanzejuyouxiangtongzazhongyoushiquxiangdeyouliangzijiaoxi,fangzhigailiangleyuanyoudebuliangxingzhuangyoudailailexindebuliangxingzhuang,yijizaochengpeihelixiajiangdengwentifasheng。huijiaohoudaiyeyinggaijinxingpeiheliceding,zaibutongdezimeixizhongxuanzepeiheligaodeyingyong。
三、玉米雜交種的選育
雜交種的選育就是將自交係或其他優良親本材料組配成雜交組合,通過測交鑒定、品種比較和各類品種比較試驗選育出符合育種目標要求的強優勢雜交種。
1.雜za交jiao種zhong的de自zi交jiao係xi選xuan育yu。自zi交jiao係xi的de選xuan育yu已yi經jing為wei雜za交jiao種zhong選xuan育yu提ti供gong了le大da量liang資zi料liao和he信xin息xi。在zai應ying用yong一yi組zu優you良liang自zi交jiao係xi選xuan育yu雜za交jiao種zhong之zhi前qian,需xu要yao對dui所suo使shi用yong的de親qin本ben有you較jiao詳xiang細xi的de了le解jie,並bing遵zun照zhao某mou些xie原yuan則ze在zai親qin本ben間jian組zu配pei:親緣關係遠,地理差異大,例如采用國外係x國內係的組配模式已經被國內育種家廣泛認識;leixingyouchayi,xingzhuangyaohubu,leixingjiandechayitongchangfanyinglejiyinxingdechayi,xingzhuanghubugengjiazhongyao,tebieshizhongyaodekangxingxingzhuang,zhishaozaiqinbenzhiyizhongyaocunzai;農藝性狀和種子生產性能好,這對於新雜交種的應用極為重要;配合力要高,這是最基本的條件之一,特別是產量性狀的配合力要高。
2.雜交種的選育技術簡單。單交種的組配除了育種者根據自己的經驗選配親本之外,通常采用最多的方法有兩種:一是輪配法:單交組合數=P(P-1)/2,其中的P為親本數;二是骨幹係法:單交組合數=M×N,即將自交係分為M、N兩組。在組間做可能的雜交,其中一組為骨幹自交係組成,另一組為新自交係組成。這種方法通常最為有效。http://www.98fo.cn



